那位高手能帮我分析一下这句话的成分,还有这是什么句型啊!

Immediately relevant to game theory are the sex ratios in certain parasitic wasp species that have a large excess of females.
与博弈理论更为直接相关的是某些寄生蜂物种的性别比例,该物种拥有数量过多的雌蜂.
这句话的主干其实是简单句--主系表结构.
Immediately relevant to game theory充当主语；
are系动词；
the sex ratios in certain parasitic wasp species表语,其后带一个定语从句 that have a large excess of females,修饰前面的parasitic wasp species,关系代词that在从句中充当主语.